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Numerous genes encoding toxic chemicals including proteinaceous inhibitors, lectins and various enzymes that participate in the synthesis of secondary metabolites are induced in resistant plants after Hessian fly attack 21,24,25

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While apoptosis is induced by p53 through activating pro-apoptotic genes, levels of p53 increase by silymarin treatment in a dose dependently manner which leads to cytochrome C release, activating many pro-apoptotic genes such as APAF-1 and caspase-9